Cod is the common name for the demersal fish genus Gadus, belonging to the family Gadidae. Cod is also used as part of the common name for a number of other fish species, and some species suggested to belong to genus Gadus are not called cod (the Alaska pollock).
The two most common species of cod are the Atlantic cod (Gadus morhua), which lives in the colder waters and deeper sea regions throughout the North Atlantic, and the Pacific cod (Gadus macrocephalus), found in both eastern and western regions of the northern Pacific. Gadus morhua was named by Linnaeus in 1758. (However, G. morhua callarias, a low-salinity, nonmigratory race restricted to parts of the Baltic, was originally described as Gadus callarias by Linnaeus.)
Cod is popular as a food with a mild flavour and a dense, flaky, white flesh. Cod livers are processed to make cod liver oil, an important source of vitamin A, vitamin D, vitamin E, and omega-3 fatty acids (EPA and DHA). Young Atlantic cod or haddock prepared in strips for cooking is called scrod. In the United Kingdom, Atlantic cod is one of the most common ingredients in fish and chips, along with haddock and plaice.
- 1 Species
- 2 Characteristics
- 3 Distribution
- 4 Life cycle
- 5 Ecology
- 6 Fisheries
- 7 Aquaculture
- 8 As food
- 9 History
- 10 1990's East Canadian cod collapse and management
- 11 See also
- 12 References
- 13 Further reading
- 14 External links
At various times in the past, taxonomists included many species in the genus Gadus. Most of these are now either classified in other genera, or have been recognized as simply forms of one of three species. All these species have a number of common names, most of them ending with the word "cod", whereas other species, as closely related, have other common names (such as pollock and haddock). However, many other, unrelated species also have common names ending with cod. The usage often changes with different localities and at different times.
Cod in the genus Gadus
Three species in the genus Gadus are currently called cod:
|Cod in the genus Gadus:|
|Common name||Scientific name||Maximum
|Atlantic cod||Gadus morhua Linnaeus, 1758||200 cm||100 cm||96.0 kg||25 years||4.4||||||||Vulnerable|
|Pacific cod||Gadus macrocephalus Tilesius, 1810||119 cm||cm||22.7 kg||18 years||4.0||||||||Not assessed|
|Greenland cod||Gadus ogac Richardson, 1836||77.0 cm||cm||kg||12 years||3.6||||||||Not assessed|
Cod forms part of the common name of many other fish no longer classified in the genus Gadus. Many are members of the family Gadidae; others are members of three related families within the order Gadiformes whose names include the word "cod": the morid cods, Moridae (100 or so species); the eel cods, Muraenolepididae (four species); and the Eucla cod, Euclichthyidae (one species). The tadpole cod family (Ranicipitidae) has now been placed in Gadidae.
|Common name||Scientific name||Maximum
|Arctic cod||Arctogadus glacialis (Peters, 1872)||32.5 cm||cm||kg||years||3.8||||||Not assessed|
|East Siberian cod||Arctogadus borisovi Dryagin, 1932||55.6 cm||cm||1.5 kg||years||3.9||||||Not assessed|
|Eucla cod||Euclichthys polynemus McCulloch, 1926||35.0 cm||22.5 cm||kg||years||3.6||||||Not assessed|
|Common ling||Molva molva (Linnaeus, 1758)||200 cm||106 cm||45.0 kg||25 years||4.3||||||||Not assessed|
|Pelagic cod||Melanonus gracilis Günther, 1878||18.7 cm||cm||kg||years||3.5||||||Not assessed|
|Polar cod||Boreogadus saida (Lepechin, 1774)||40.0 cm||25.0 cm||kg||7 years||3.1||||||||Not assessed|
|Poor cod||Trisopterus minutus (Linnaeus, 1758)||40.0 cm||20.0 cm||kg||5 years||3.8||||||Not assessed|
|Rock cod||Lotella rhacina (Forster, 1801)||50.0 cm||cm||kg||years||3.5||||||Not assessed|
|Saffron cod||Eleginus gracilis (Tilesius, 1810)||55.0 cm||cm||1.3 kg||15 years||4.1||||||||Not assessed|
|Small-headed cod||Lepidion microcephalus Cowper, 1956||48.0 cm||cm||kg||years||3.5||||||Not assessed|
|Tadpole cod||Guttigadus globosus (Paulin, 1986)||18.1 cm||cm||kg||3.5 years||||||Not assessed|
Some fish commonly known as cod are unrelated to Gadus. Part of this name confusion is market-driven. Severely shrunken Atlantic cod stocks have led to the marketing of cod replacements using culinary names of the form "x cod", according to culinary rather than phyletic similarity. The common names for the following species have become well established; note that all inhabit the Southern Hemisphere.
- Blue cod Parapercis colias
- Eastern freshwater cod Maccullochella ikei
- Mary River cod Maccullochella peelii mariensis
- Murray cod Maccullochella peelii peelii
- Potato cod Epinephelus tukula
- Sleepy cod Oxyeleotris lineolatus
- Trout cod Maccullochella macquariensis
- The notothen family, Nototheniidae, including:
- Rock cod, reef cod, and coral cod
Almost all coral cod, reef cod or rock cod are also in order Perciformes. Most are better known as groupers, and belong to the family Serranidae. Others belong to the Nototheniidiae. Two exceptions are the Australasian red rock cod, which belongs to a different order (see below), and the fish known simply as the rock cod and as soft cod in New Zealand, Lotella rhacina, which as noted above actually is related to the true cod (it is a morid cod).
From the order Scorpaeniformes:
Marketed as cod
Some fish that do not have "cod" in their names are sometimes sold as cod. Haddock and whiting belong to the same family, the Gadidae, as cod.
Cods of the genus Gadus have three rounded dorsal and two anal fins. The pelvic fins are small, with the first ray extended, and are set under the gill cover (i.e. the throat region), in front of the pectoral fins. The upper jaw extends over the lower jaw, which has a well-developed chin barbel. The eyes are medium-sized, approximately the same as the length of the chin barbel. Cod have a distinct white lateral line running from the gill slit above the pectoral fin, to the base of the caudal or tail fin. The back tends to be a greenish to sandy brown, and shows extensive mottling, especially towards the lighter sides and white belly. Dark brown colouration of the back and sides is not uncommon, especially for individuals that have resided in rocky inshore regions.
The Atlantic cod can change colour at certain water depths. It has two distinct colour phases: gray-green and reddish brown. Its average weight is 5–12 kilograms (11–26 lb), but specimens weighing up to 100 kilograms (220 lb) have been recorded. Pacific cod are smaller than Atlantic cod and are darker in colour.
Atlantic cod (Gadus morhua) live in the colder waters and deeper sea regions throughout the North Atlantic. Pacific cod (Gadus macrocephalus) is found in both eastern and western regions of the Pacific.
Atlantic cod divide into several stocks, including the Arcto-Norwegian, North Sea, Faroe, Iceland, East Greenland, West Greenland, Newfoundland, and Labrador stocks. There seems to be little interchange between the stocks, although migrations to their individual breeding grounds may involve distances of 200 miles (320 km) or more.
Atlantic cod occupy varied habitat, favouring rough ground, especially inshore, and are demersal in depths between 20 and 200 feet (6.1 and 61.0 m), 80 metres (260 ft) on average, although not uncommonly to depths of 600 metres (2,000 ft). Off the Norwegian and New England coasts and on the Grand Banks of Newfoundland, cod congregate at certain seasons in water of 30–70 metres (98–230 ft) depth. Cod are gregarious and form schools, although shoaling tends to be a feature of the spawning season.
Spawning of northeastern Atlantic cod occurs between January and April (March and April are the peak months), at a depth of 200 metres (660 ft) in specific spawning grounds at water temperatures between 4 and 6 °C (39 and 43 °F). Around the UK, the major spawning grounds are in the middle to southern North Sea, the start of the Bristol Channel (north of Newquay), the Irish Channel (both east and west of the Isle of Man), around Stornoway, and east of Helmsdale.
Prespawning courtship involves fin displays and male grunting, which leads to pairing. The male inverts himself beneath the female, and the pair swim in circles while spawning. The eggs are planktonic and hatch between eight and 23 days, with larva reaching 4 millimetres (0.16 in) in length. This planktonic phase lasts some ten weeks, enabling the young cod to increase its body weight by 40-fold, and growing to about 2 centimetres (0.79 in). The young cod then move to the seabed and change their diet to small benthic crustaceans, such as isopods and small crabs. They increase in size to 8 centimetres (3.1 in) in the first six months, 14–18 centimetres (5.5–7.1 in) by the end of their first year, and to 25–35 centimetres (9.8–13.8 in) by the end of the second. Growth tends to be less at higher latitudes. Cod reach maturity at about 50 centimetres (20 in) at about 3 to 4 years of age.
In the Baltic Sea the most important prey species are herring and sprat. Many studies that analyze the stomach contents of these fish indicate that cod is the top predator, preying on the herring and sprat. Sprat form particularly high concentrations in the Bornholm Basin in the southern Baltic Sea. Although cod feed primarily on adult sprat, sprat tend to prey on the cod eggs and larvae.
Cod and related species are plagued by parasites. For example, the cod worm, Lernaeocera branchialis, starts life as a copepod-like larva, a small free-swimming crustacean. The first host used by the larva is a flatfish or lumpsucker, which it captures with grasping hooks at the front of its body. It penetrates the fish with a thin filament, which it uses to suck the fish's blood. The nourished larvae then mate on the fish. The female larva, with her now fertilized eggs, then finds a cod, or a cod-like fish such as a haddock or whiting. There the larva clings to the gills while it metamorphoses into a plump sinusoidal wormlike body with a coiled mass of egg strings at the rear. The front part of the worm's body penetrates the body of the cod until it enters the rear bulb of the host's heart. There, firmly rooted in the cod's circulatory system, the front part of the parasite develops like the branches of a tree, reaching into the main artery. In this way, the worm extracts nutrients from the cod's blood, remaining safely tucked beneath the cod's gill cover until it releases a new generation of offspring into the water.
The 2006 northwest Atlantic cod quota is 23,000 tons, representing half the available stocks, while the northeast Atlantic quota is 473,000 tons. Pacific cod is currently enjoying strong global demand. The 2006 total allowable catch (TAC) for the Gulf of Alaska and Aleutian Islands was 260,000 tons.
Farming of Atlantic cod has received a significant amount of interest due to the overall trend of increasing cod prices alongside reduced wild catches. However, progress in creating large scale farming of cod has been slow, mainly due to bottlenecks in the larval production stage, where survival and growth are often unpredictable. It has been suggested that this bottleneck may be overcome by ensuring cod larvae are fed diets with similar nutritional content as the copepods they feed on in the wild  Recent examples have shown that increasing dietary levels of minerals such as selenium, iodine and zinc may improve survival and/or biomarkers for health in aquaculture reared cod larvae.
Cod is popular as a food with a mild flavor and a dense, flaky white flesh. Cod livers are processed to make cod liver oil, an important source of vitamin A, vitamin D, vitamin E and omega-3 fatty acids (EPA and DHA).
Young Atlantic cod or haddock prepared in strips for cooking is called scrod. In the United Kingdom, Atlantic cod is one of the most common ingredients in fish and chips, along with haddock and plaice. Cod's soft liver can be tinned (canned) and eaten. Cod is mainly consumed in Portugal, Spain, Italy and Brazil.
Cod has been an important economic commodity in international markets since the Viking period (around 800 AD). Norwegians travelled with dried cod and soon a dried cod market developed in southern Europe. This market has lasted for more than 1,000 years, enduring the Black Death, wars and other crises, and is still an important Norwegian fish trade. The Portuguese began fishing cod in the 15th century. Clipfish is widely enjoyed in Portugal. The Basques played an important role in the cod trade, and allegedly found the Canadian fishing banks before Columbus' discovery of America. The North American east coast developed in part due to the vast cod stocks. Many cities in the New England area are located near cod fishing grounds. The fish was so important to the history and development of Massachusetts, the state's House of Representatives hung a wood carving of a codfish, known as the Sacred Cod of Massachusetts, in its chambers.
Apart from the long history, cod differ from most fish because the fishing grounds are far from population centers. The large cod fisheries along the coast of North Norway (and in particular close to the Lofoten islands) have been developed almost uniquely for export, depending on sea transport of stockfish over large distances. Since the introduction of salt, dried and salted cod (clipfish or 'klippfisk' in Norwegian) has also been exported. By the end of the 14th century, the Hanseatic League dominated trade operations and sea transport, with Bergen as the most important port.
In the 17th and 18th centuries in the New World, especially in Massachusetts and Newfoundland, cod became a major commodity, creating trade networks and cross-cultural exchanges. In 1733, Britain tried to gain control over trade between New England and the British Caribbean by imposing the Molasses Act, which they believed would eliminate the trade by making it unprofitable. The cod trade grew instead, because the "French were eager to work with the New Englanders in a lucrative contraband arrangement". In addition to increasing trade, the New England settlers organized into a "codfish aristocracy". The colonists rose up against Britain's "tariff on an import".
In the 20th century, Iceland re-emerged as a fishing power and entered the Cod Wars. In the late 20th and early 21st centuries, fishing off the European and American coasts severely depleted stocks and become a major political issue. The necessity of restricting catches to allow stocks to recover upset the fishing industry and politicians are reluctant to hurt employment.
1990's East Canadian cod collapse and management
- For information, see Collapse of the Atlantic northwest cod fishery.
- Harold Innis and the cod fishery, for the Canadian industry
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- Kamisaka, Y.; Jordal, A.E.O.; Edvardsen, R.B.; Kryvi, H.; Otterlei, E.; Rønnestad, I (2010). "A case report on the distended gut syndrome (DGS) in cultured larvae of Atlantic cod (Gadus morhua)". Aquaculture. 309 (1–4): 38–48. doi:10.1016/j.aquaculture.2010.09.006.
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- Hamre, K.; Srivastava, A.; Rønnestad, I.; Mangor-Jensen, A.; Stoss, J (2008). "Several micronutrients in the rotifer Brachionus sp. may not fulfil the nutritional requirements of marine fish larvae". Aquaculture Nutrition. 14: 51–60. doi:10.1111/j.1365-2095.2007.00504.x.
- Hamre, K.; Mollan, T.A.; Sæle, Ø.; Erstad, B (2008). "Rotifers enriched with iodine and selenium increase survival in Atlantic cod (Gadus morhua) larvae". Aquaculture. 284 (1–4): 190–195. doi:10.1016/j.aquaculture.2008.07.052.
- Penglase, S.; Nordgreen, A.; van der Meeren, T.; Olsvik, P.; Sæle, O.; Baeverfjord, G.; Helland, S.; Hamre, K (2010). "Increasing the level of selenium in rotifers (Brachionus plicatilis 'Cayman') enhances the mRNA expression and activity of glutathione peroxidase in Atlantic cod (Gadus morhua L.) larvae". Aquaculture. 306 (1–4): 259–269. doi:10.1016/j.aquaculture.2010.05.011.
- Penglase, S.; Harboe, T.; Sæle, O.; Helland, S.; Nordgreen, A.; Hamre, K (2013). "Iodine nutrition and toxicity in Atlantic cod (Gadus morhua) larvae". PeerJ. 1:e20: e20. doi:10.7717/peerj.20.
- Penglase, S.; Hamre, K.; Olsvik, P.; Grøtan, E.; Nordgreen, A (2013). "Rotifers enriched with iodine, copper and manganese had no effect on larval cod (Gadus morhua) growth, mineral status or redox system gene mRNA levels". Aquaculture Research. 46 (8): 1793. doi:10.1111/are.12332.
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|Presentation by Mark Kurlansky on Cod: A Biography of the Fish That Changed the World, August 15, 1998, C-SPAN|
- Dean L.Y. Bavington. Managed Annihilation: An Unnatural History of the Newfoundland Cod Collapse (University of British Columbia Press; 2010) 224 pages. Links the collapse of Newfoundland and Labrador cod fishing to state management of the resource.
- Mark Kurlansky. Cod: A Biography of the Fish That Changed the World (1997)
- Cobb, John N (1916). Pacific Cod Fisheries. Bureau of Fisheries Document. no. 830. Washington, DC: Government Printing Office. OCLC 14263968.
- Shields, Edward (2001). Salt of the sea: the Pacific Coast cod fishery and the last days of sail. Lopez Island, Wash.: Heritage House. ISBN 978-1-894384-35-3.
- Four Fish, by Paul Greenberg, ISBN 978-1-59420-256-8
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